Publication Companion
bioRxiv preprint posted January 12, 2026
Turning Dynamics Resource

Speed-Dependent Turning Strategies in Quadrupedal Locomotion

Interactive supplement and reproducibility hub for the bioRxiv preprint Speed-Dependent Turning Strategies in Quadrupedal Locomotion: Insights from Computational Modeling. The paper extends a previously published quadrupedal locomotion model to compare three turning asymmetries, measure turning curvature and inner/outer duty-factor ratios across walking speeds, and test how axial foot-placement shifts expand the stable turning range.

Yaroslav I. Molkov, Mohammed A. Y. Mohammed, Tommy Stell, Amelia Harralson, Russell Jeter, and Ilya A. Rybak

Research group: Yaroslav Molkov's group at Georgia State University

bioRxiv DOI 10.64898/2026.01.12.699101 Interactive animator included
Figure 7 From The Preprint
Figure 7 from the preprint comparing maximal stable curvature across body bending, lateral force, and lateral limb shift

Figure 7 compares the maximum stable curvature produced by body bending, lateral force, and lateral limb shift across walking speeds under the strategy-specific axial-shift settings used in the paper.

Key Result

Peak stable turning strategy depends on walking speed

Condensed from Figures 7, 9, and 10 of the preprint
Low speed

Body bending yields the highest stable curvature at low speed

With axial shift used as stride control, body bending is the strongest low-velocity strategy, especially below about 5 cm/s, where axial flexion can reorient the body while remaining statically stable.

Medium speed

Lateral force reaches the largest curvature at intermediate speed

At roughly 5 to 15 cm/s, forelimb-applied lateral force produces the largest maximum stable curvature in the model, outperforming body bending once turning demands become more dynamic.

High speed

Lateral limb shift performs best at higher walking speeds

At higher speeds, the best-performing strategy is a mediolateral shift of limb touchdown positions. In the model, that foot-placement asymmetry preserves maneuverability as roll-over risk grows.

Coordination

Forelimb steering asymmetry is consistent, hindlimb timing is not

The paper tracks inner/outer duty-factor ratios and finds a consistent steering-related asymmetry in the forelimbs across strategies, while hindlimb duty factors reorganize differently for body bending, lateral force, and lateral limb shift.

Paper Context

What this companion site is for

Study summary

The paper extends the Molkov et al. (2024) quadrupedal locomotion model to compare three turning asymmetries: body bending (geometric), lateral force applied to the forelimbs (dynamic), and lateral limb shift at touchdown (kinematic). Across walking speeds, the authors evaluate the maximum curvature that can be sustained during stable locomotion and track how turning reshapes inner/outer duty-factor ratios.

A central modeling result is that axial (sagittal) shift of the footfall targets acts as stride control: it enlarges the stable turning envelope for all three strategies, but in strategy-specific ways. Under those optimized conditions, body bending performs best at low speed, lateral force at intermediate speed, and lateral limb shift at higher speed. The paper also tests pairwise combinations, showing that mixed strategies mainly bridge the transition zones between the single-strategy speed regimes.

Citation

Molkov YI, Mohammed MAY, Stell T, Harralson A, Jeter R, Rybak IA. Speed-Dependent Turning Strategies in Quadrupedal Locomotion: Insights from Computational Modeling. bioRxiv (2026). doi:10.64898/2026.01.12.699101
Companion Materials

Move between the paper, the tool, and the code

Interactive Animator

Explore steering controls in the browser and inspect the turning behaviors without building video files first. This is the main interactive entry point for the companion site.

Launch animator

Preprint and PDF

Read the full paper on bioRxiv or jump directly to the PDF when you want the publication itself rather than the interactive supplement.

Open preprint

Source and Reproducibility

Download the project archive containing the C++ model, scripts, and build targets used to regenerate the example media and parameter sweeps.

Download source
Reproducibility

Build locally and regenerate outputs

Local reproduction requires g++, make, gnuplot, ffmpeg, and bc. The commands below match the project targets used to build the principal example animations and parameter sweeps.

Install and build

sudo apt update
sudo apt install build-essential gnuplot ffmpeg bc
make

Generate example outputs

make results/lateral_force.mp4
make results/body_bending.mp4
make results/lateral_shift.mp4
make delta
make force
make shift